Coleoptera
Staphylinidae -Scaphidinae
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http://www.nhm.ku.edu/ksem/projects/scaphcat.htm

World Catalogue of Scaphidiinae
(Coleoptera: Staphylinidae)

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Richard A. B. Leschen
Department of Entomology, Michigan State University, East Lansing, MI, 48824
E-mail: leschen@pilot.msu.edu
AND
Ivan Löbl
Muséum d'Histoire naturelle, Route de Malagnou 1, case postale 6434, CH-1211 Genčve 6, Switzerland
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Introduction
The subfamily Scaphidiinae is a member of the Oxyteline Group of Staphylinidae (Lawrence and Newton 1982). Unlike most members of the Oxyteline Group, and the majority of Staphylinidae, scaphidiines do not have the flexible staphylinoid body form, but are box-like and highly convex, with the elytra covering the abdomen. Some members, such as Diatelium wallacei Pascoe, with its greatly elongate neck and short body, is unlike any other Coleoptera. Based on larval characters Kasule (1966) was the first to recognize Scaphidiinae as a subfamily of Staphylinidae, despite their long taxonomic history set apart as separate families. It has been a slow process for coleopterists to accept the firm placement of Scaphidiinae within staphylinids, and many works published after Kasule's (1966) paper treat scaphidiines as a separate family (see review in Leschen and Löbl 1995), despite numerous adult characters that support the correct familial placement (Lawrence and Newton 1982, Newton and Thayer 1992).

The subfamily Scaphidiinae contains approximately 1300 described species worldwide (Löbl 1992) and is unequivocally monophyletic (Leschen and Löbl 1995), although its direct sister relationships are uncertain within the Oxyteline Group. It is currently divided into four tribes, with Scaphisomatini having the highest number of described species (967, Table 1) and the most enigmatic phylogenetic relationships. Leschen and Löbl (1995) studied the phylogenetic relationships among the scaphidiine tribes, and the genera contained in Cypariini, Scaphiini, and Scaphidiini demonstrating the monophyly of Scaphisomatini. Scaphisomatini includes five subtribes (Table 1), reflecting higher categories defined by Achard (1924). Preliminary phylogenetic analyses demonstrate that each of these subtribes appears polyphyletic (Leschen and Löbl unpublished).

The best collecting methods for scaphidiines include flight intercept traps, sifting leaf litter, and hand collection from fungi. Specimens are rare in collections from forest canopies, suggesting that scaphidiines are limited to understory habitats. Mycophagy is a somewhat unique feature among the largely saprophagous and predaceous Oxyteline Group and is present among all members of Scaphidiinae (Lawrence & Newton 1982, Newton 1984). Associated with this shift to mycophagy is a group of morphological and behavioral adaptations that include novel mouthpart structures and canopy-retreat building behaviors in larvae not present anywhere else in Coleoptera (Newton 1984, 1991; Lawrence 1988; Leschen 1993, 1994; Hanley 1996). In general, adults occur on a variety of host fungi; in contrast, larvae are more restricted in their occurrence on fungal hosts (Newton 1984). Many species of Baeocera and Scaphobaeocera are specialists on slime molds (Lawrence and Newton 1980, Newton & Stephenson 1990), while members of Scaphisoma and Caryoscapha occur on relatively persistent wood-rotting fungi (Newton 1984; Ashe 1984; Leschen 1988). Species of Toxidium (Scaphisomatini), and other taxa contain species associated with fungal growth in leaf-litter and under rotten logs. Members of the subtribe Baeoceridiina are associated with Old World termites that build fungal gardens.

Table 1. Scaphisomatini subtribes, genera, number of described species and distribution.
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Baeoceridiina Achard
Antongilium Pic; 25 species; Africa
Baeoceridium Reitter; 4 species; Africa,Asia
Collartium Pic; 1 species; Africa
Mordelloscaphium Pic; 1 species; Africa,Borneo
Mysthrix Champion; 2 species; Asia,Sumatra
Sapitia Achard; 4 species; Asia
Termitoscaphium Löbl; 1 species; Sulawesi
Termitoxidium Pic; 1 species; Africa
Trichoscaphella Reitter; 1 species; Africa
Vituratella Reitter; 2 species; Africa,Madagascar
Beaocerina Achard
Alexidia Reitter; 1 species; South America
Amalocera Erichson; 5 species; South America
Amaloceromorpha Pic; 1 species; Burma

Baeocera Erichson; 217 species; World wide
Brachynoposoma Löbl; 2 species; New Caledonia
Brachynopus Broun; 1 species; New Zealand
Nesoscapha Vinson; 2 species; Mauritius
Pseudobironiella Löbl 3 species; Madagascar
Pseudobironium Pic; 25 species; Asia,New Caledonia
Scaphischema Reitter; 1 species; Mediterranean
Zinda Löbl 2 species; New Caledonia
Heteroscaphina Achard

Birocera Löbl 2 species; Sulawesi
Bironium Csiki; 27 species; Asia,New Guinea
Scaphisomatina Casey

Caryoscapha Ganglbauer; 4 species; Holarctic
Curtoscaphosoma Pic; 1 species; Africa
Kathetopodion Löbl 1 species; Borneo
Macroscaphosoma Löbl 2 species; Africa
Metalloscapha Löbl 1 species; New Guinea
Scaphisoma Leach; 547 species; World wide
Toxidiina Achard

Baeotoxidium Löbl 7 species; Asia
Paratoxidium Vinson; 1 species; Mauritius
Scaphicoma Motschulsky; 11 species; Africa,Asia
Scaphobaeocera Csiki; 60 species; Old World
Scaphoxium Löbl 26 species; Old World
Toxidium LeConte; 40 species; World wide
Unplaced

Afroscaphium Löbl 2 species; Africa
Xotidium Löbl 4 species; Asia

The catalogue provided here contains all of the species in the tribes Scaphiini, Scaphidiini, and Cypariini. Some taxonomic changes made by Leschen and Löbl (1995) are included in the list and this paper should be consulted regarding details about current classification and phylogeny. The genera of Scaphisomatini are given in Table 1. Currently Ivan is working on a complete species catalogue with complete citations, type specimen depositions and other useful information that will be added to this catalogue when available. Other work in progress is a cladistic analysis of Scaphisomatini which will identify monophyletic subtribes and genera and provide the basis for a natural classification of the tribe. We are continuing to pursue grant money to support our on-going studies of Scaphidiinae systematics and evolution.


SPECIES CATALOGUE

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CYPARIINI Achard, 1924
Cyparium Erichson, 1845
anale Reitter, 1880; Neotropical
ater Casey, 1900; Nearctic
basilewskyi Pic, 1955; Afrotropical
bowringi Achard, 1922; SE Asia
championi Matthews, 1888; Neotropical
collare Pic, 1920; Neotropical
concolor (Fabricius, 1801); Nearctic
flavipes LeConte, 1860
substriatum Reitter, 1880
ferrugineum Pic, 1920; Neotropical
flavosignatum Zayas, 1989; Neotropical
ssp. bicolorum Zayas, 1989
ssp. funebre Zayas, 1989
ssp. splendidum Zayas, 1989
formosanum Miwa & Mitono, 1943; SE Asia
grilloi Pic, 1920; Neotropical
grouvellei Pic, 1920; Neotropical
humerale Achard, 1922; Neotropical
inclinans Kirsch, 1873; Neotropical
javanum Löbl, 1990; SE Asia
khasianum Löbl, 1984; SE Asia
laevisternale Nakane, 1956; E Palaearctic
mathami R. Oberthür, 1884; Neotropical
mikado Achard, 1923; E Palaearctic
minutum Pic, 1931; Neotropical
montanum Achard, 1922; SE Asia
monticola Miwa & Mitono, 1943; SE Asia
multistriatum Pic, 1954; Africa
nigronotatum Pic, 1931; Neotropical
oberthueri Pic, 1955; Neotropical
palliatum Erichson, 1845; Neotropical
pallidum Pic, 1955; Africa
peruvianum Pic, 1947; Neotropical
piceum Reitter, 1880; S Africa
plagipenne Achard, 1922; SE Asia
punctatum Pic, 1916; SE Asia
pygidiale Achard,1922; Neotropical
ruficolle Achard, 1922; Neotropical
rufohumerale Pic, 1931; Neotropical
rufonotatum Pic, 1916; Neotropical
sallei Matthews, 1888; Neotropical
semirufum Pic, 1917; SE Asia
siamense Löbl, 1990; SE Asia
sibiricum Solsky, 1871; E Palaearctic
submetallicum Reitter, 1880; ? SE Asia
tamil Löbl, 1979; SE Asia
terminale Matthews, 1888; Neotropical
tenenbaumi Pic, 1926; E Palaearctic
testaceicorne Pic, 1931; Neotropical
testaceum Pic, 1920; SE Asia
variabile Pic, 1955; Afrotropical
var. atrocinctum Pic, 1955
var. diversipenne Pic, 1955
variegatum Achard, 1920; SE Asia
yunnanum (Achard, 1920); SE Asia

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SCAPHIINI Achard, 1924
Ascaphium Lewis, 1893
apicale Lewis, 1893; E Palaearctic
minor Pic, 1954; SE Asia
ochripes Löbl, 1992; SE Asia
sinense Pic, 1954; SE Asia
sulcipenne Lewis, 1893; E Palaearctic
tibiale Lewis, 1893; E Palaearctic
tonkinense Achard, 1921; SE Asia
Episcaphium Lewis, 1893

*tranferred from Phenoscaphium by Leschen and Löbl (1995)

callosipenne (Achard, 1922)*; SE Asia
saucineum (Motschulsky, 1859); SE Asia
semirufum Lewis, 1893; E Palaearctic
var. ruficolle Lewis, 1893
unicolor Löbl, 1992; SE Asia
Scaphium Kirby, 1837

castanipes Kirby, 1837; Nearctic
ferrugineum Reitter, 1880; ?S Africa
immaculatum (Olivier, 1790); W Palaearctic
rufipes Reitter, 1883
quadraticolle Solsky, 1874; C Asia


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SCAPHIDIINI Latrielle, 1807
Euscaphidium Achard, 1922
tuberosum Achard, 1922; SE Asia
Scaphidium Olivier, 1790

The following genera were transferred by Leschen and Löbl (1995): * =3D Scaphidiolum Achard, 1922; ** =3DHemiscaphium Achard, 1922; *** =3D Ascaphidium Pic, 1915; + =3D Scaphidiopsis Achard, 1922; ++ =3D =46alsoascaphidium Pic, 1923; +++ =3D Cribroscaphium Pic, 1920

abdominale Achard, 1920; SE Asia
abyssinicum (Pic,,1954)*; Africa
alpicola Blackburn, 1891; Australia
alpicolum Blackburn
alternans Löbl, 1978; New Guinea
amurense Solsky,1871; E Palaearctic
ab. bodemeyeri Reitter, 1913
andrewesi Achard, 1922); SE Asia
var. femoratum Achard, 1922
angolense Pic, 1940; Africa
angustatum Pic, 1920; Madagascar
antennatum Reitter, 1880; Nearctic
anthrax Achard, 1920; SE Asia
apicicorne (Pic,1953)*; Madagascar
arrowi Achard, 1920; SE Asia
aterrimum Reitter, 1880; SE Asia
atricolor Pic, 1915; SE Asia
atripenne Gestro, 1879; SE Asia
atripes Pic 1946; Africa
atrosuturale Pic, 1915; SE Asia
atrum Matthews, 1888; Neotropical
baconi Pic, 1915; SE Asia
var.assamense Pic, 1915
var. multimaculatum Pic, 1915
var. semifasciatum Pic, 1915
var. uniplagatum Achard, 1922
badium Heller, 1917; SE Asia
basale Cas telnau, 1940*; Madagascar
basilewskyi (Pic, 1955)*; Africa
benitense (Achard, 1922)*; Africa
bicinctum Achard, 1920; SE Asia
anamalaiense Löbl, 1971
nathani Löbl, 1971
bicolor Castelnau, 1840; Madagascar
var. unicolor Castelnau, 1840
bifasciatum Pic, 1915; SE Asia
bilineatithorax (Pic, 1931)*; SE Asia
binhanum (Pic, 1925)*; SE Asia
binigronotatum ( Pic, 1931)*; Africa
binominatum Achard, 1915; SE Asia
notaticolle Pic, 1915
binotatum Achard 1915; SE Asia
bipunctatum Redtenbacher, 1867; Neotropical
bisbimaculatum Pic, 1917; Neotropical
biseriatum Champion, 1927; SE Asia
biundulatum Champion, 1927; SE Asia
bolivianum (Pic, 1931); Neotropical
borneense Pic, 1915**; SE Asia
brunneopictum (Achard, 1922)**; SE Asia
carinense Achard, 1920; SE Asia
castaneicolor Csiki, 1924; SE Asia
castaneum Perty, 1830; Neotropical
celebense Pic, 1915; SE Asia
cerasinum R. Oberth=FCr, 1884; Neotropical
chapuisi Gestro, 1880; SE Asia
cheesmanae Löbl, 1975; New Guinea
chujoi Löbl, 1967; E Palaearctic
cinnamoneum Champion, 1927; SE Asia
clathratum Achard, 1920; SE Asia
coerulans Löbl, 1979; New Guinea
comes Löbl, 1968; E Palaearctic
compressum Achard, 1915; SE Asia
confusum (Pic, 1926)*; SE Asia
conjunctum Motschulsky, 1859; SE Asia
consimile Achard, 1920; SE Asia
consobrinum Castelnau, 1840; Madagascar
coomani (Pic, 1925); SE Asia
coronatum Reitter, 1880; Australia
cyanellum R. Oberth=FCr, 1884; SE Asia
cyanipenne Gestro, 1879; New Guinea
decorsei Achard, 1920; Madagascar
delatouchei Achard, 1920; SE Asia
deletum Heer, 1847; (fossil) Tertiary, Oeningen
discerptum (Achard, 1922)**; SE Asia
disclusum (Achard, 19)*; SE Asia
discoidale Pic, 1920; Madagascar
disconotatum Pic, 1915; SE Asia
distinctum Achard, 1916; Australia
dohertyi Pic, 1915**; SE Asia
donckieri Pic, 1917; Madagascar
dureli (Achard, 1922); SE Asia
egregium Achard, 1922; SE Asia
atripenne Pic, 1922
elisabethae (Pic, 1954)*; Africa
ellenbergeri (Paulian, 1951)*; Africa
elongatum Achard, 1915; SE Asia
emarginatum Lewis, 1893; E Palaearctic
eraserheadi Leschen & Löbl, 1995; Africa
exclamans R. Oberth=FCr, 1884; Neotropical
exornatum R. Oberth=FCr, 1884; Australia
australe Achard, 1916
fainanense Pic, 1915; SE Asia
fairmairei Pic, 1920; Madagascar
fasciatomaculatum R. Oberth=FCr, 1884; Neotropical
fasciatum Castelnau, 1840; Madagascar
fascipenne Reitter, 1880; Neotropical
feai Pic, 1920; SE Asia
femorale Lewis, 1893; E Palaearctic
flavofasciatum Champion, 1913; Neotropical
flavomaculatum Miwa & Mitono, 1943; SE Asia
formosanum Pic, 1915; SE Asia
fossulatum Pic, 1921; SE Asia
fraternum Achard, 1920; SE Asia
fryi Achard, 1920; SE Asia
fukienense Pic, 1954; SE Asia
gabonicum (Paulian, 1951)*; Africa
geniculatum R. Oberth=FCr, 1884; Neotropical
gestroi Pic, 1920; SE Asia
gibbosum Pic, 1915; SE Asia
gounellei Pic, 1920; Neotropical
gracile Achard, 1920; SE Asia
grande Gestro, 1880; SE Asia
inimpressum Pic, 1920
sunannulatum Pic, 1915
var. melanopus Achard, 1924
grandidieri Achard, 1920; Madagascar
grouvellei Achard, 1920; SE Asia
gurung Löbl, 1992; SE Asia
hageni Weyenbergh; (fossil) Jurassic, Solenhofen
harmandi Achard, 1920; SE Asia
hexaspilotum (Achard, 1924); SE Asia
holzschuhi Löbl, 1992; SE Asia
ifanense (Pic, 1947)*; Africa
impictum Boheman, 1851; S Africa
impuncticolle Pic, 1915**; SE Asia
incisum Lewis, 1893; E Palaearctic
incrassatum Achard, 1920; SE Asia
indicum Löbl, 1979; SE Asia
innotatum (Pic, 1940)*; SE Asia
inornatum Gestro, 1880; SE Asia
interruptum Fairmaire, 1897; Madagascar
irregulare Pic, 1920+++; SE Asia
jacobsoni Achard, 1921; SE Asia
japonum Reitter, 1877; E Palaearctic
japonicum Lewis, 1893
longipes Lewis, 1891
javanum Pic, 1915; SE Asia
klapperichi Pic, 1954; SE Asia
lafertei Pic, 1920; Madagascar
laosense (Pic, 1928)*; SE Asia
lateflavum (Pic, 1928)*; SE Asia
latissimum Achard, 1915; Madagascar
leleupi (Pic, 1954)*; Africa
var. atropygum Pic,1954
lesnei Achard, 1920; SE Asia
lewisi (Achard, 1923)*; E Palaearctic
lineaticolle Matthews, 1888; Neotropical
lineatipes (Pic, 1925)*; Africa
longicolle Pic, 1915; SE Asia
var. bicoloripes Pic, 1948
var. kudatense Pic, 1948
var. plagatum Achard, 1920
longipenne Achard, 1921; SE Asia
longithorax Pic, 1916; SE Asia
var. nigriventre Achard, 1924
lucidum Achard, 1915; Neotropical
lunatum Motschulsky, 1859; SE Asia
lunulatum Pic, 1921
var. bioculatum Achard, 1924
var. inconjunctum Pic, 1921
var. rufithorax Pic, 1921
luteomaculatum (Pic, 1923)*; SE Asia
luzonicum (Achard, 1924)*; SE Asia
maculaticeps (Pic, 1923)*; SE Asia
madecassum Pic, 1917; Madagascar
madurense Achard, 1915; SE Asia
malaccanum Pic, 1915; SE Asia
mangenoti (Paulian, 1951)*; Africa
marginale Reitter 1880; SE Asia
marginatum Matthews, 1888; Neotropical
martapuranum Pic, 1916; SE Asia
mastersi Macleay, 1871; (nom.dubium) Australia
matthewsi Csiki, 1904; Neotropical
unicolor Matthews, 1888
mayneui (Pic, 1954)*; Africa
medionigrum Pic, 1915; SE Asia
melanogaster Löbl, 1992; SE Asia
melli Löbl, 1972; SE Asia
metallescens Gestro, 1879; New Guinea
mexicanum Castelnau, 1840; Neotropical
minutum Pic, 1920; SE Asia
monteithi Löbl, 1976; Australia
montivagum Shir=F4zu & Morimoto, 1963; E Palaearctic
morimotoi Löbl, 1982; E Palaearctic
takashii Shir=F4zu & Morimoto, 1963
multinotatum Pic, 1921; SE Asia
multipunctatum (Pic, 1928)*; Africa
var. luluanum Pic, 1954
var. prolongatum Pic, 1954
negrito Heller, 1917; SE Asia
nepalense Löbl, 1992; SE Asia
nigripes Chevrolat, 1830; Neotropical
nigrocinctulum R. Oberth=FCr, 1884; SE Asia
nigromaculatum Reitter, 1880; SE Asia
var. effigiatum Achard, 1922
nigrosuturale Pic, 1920; SE Asia
nigrum Castelnau, 1840; Madagascar
notaticolle Pic, 1915; SE Asia
notatum (Pic, 1923); Australia
ocellatum Achard, 1920; SE Asia
var. birmanica Achard, 1920
oculare (Pic, 1923)*; Madagascar
optabile (Lewis, 1893); E Palaearctic
orbiculosum Reitter, 1880; SE Asia
ornatum Casey, 1900; Nearctic
overlaeti (Pic, 1954)*; Africa
palonense Achard, 1920; SE Asia
pantherinum R. Oberth=FCr, 1883; Neotropical
papuanum Löbl, 1975; New Guinea
pardale Castelnau, 1840+; Neotropical
var. nigripenne R. Oberth=FCr, 1884
patinoi R. Oberth=FCr, 1884; Neotropical
peninsulare Achard, 1920**; SE Asia
peraffine R. Oberth=FCr, 1884; Neotropical
perpulchrum Csiki, 1909; SE Asia
philippense Reitter, 1880; SE Asia
phungi Pic, 1923; SE Asia
picconii Gestro, 1880; SE Asia
var. sexmaculatum Reitter, 1889
piceoapicale (Pic, 1940)*; SE Asia
plagatum Achard, 1920; SE Asia
politum Fairmaire, 1899; Madagascar
prolongatum (Pic, 1928)*; Africa
pulchellum Reitter, 1880; Madagascar
punctaticolle Pic, 1923; SE Asia
punctatum Castelnau, 1840; Madagascar
punctipenne Macleay, 1871; Australia
coronatum Reitter, 1880
thoracicum Achard, 1916
pygidiale Pic, 1917; Madagascar
var. bicoloricolle Pic, 1917
quadriguttatum Say, 1823; Nearctic
amplum Cawey, 1900
piceum Melsheimer, 1846
obliteratum LeConte, 1860
quadritotatum Castelnau, 1840
quadripustulatum Say, 1823
quadrillum Fairmaire, 1898; Madagascar
var. biconjuntum Pic, 1920
quadrimaculatum Olivier, 1790; W Palaearctic
quadriplagatum Achard, 1915; SE Asia
quadriplagiatum Achard, Pic, 1916
quadripustulatum (Fabricius, 1775); Australia
bimaculatum Macleay, 1871
quinquemaculatum Pic, 1915; Neotropical
reductum (Pic, 1954)*; Africa
reitteri Lewis, 1879; E Palaearctic
insulare Achard, 1922
rosenbergi Pic, 1955; Neotropical
rouyeri Pic, 1915; SE Asia
rubicundum Reitter, 1879; Neotropical
rubricolle (Pic, 1951)*; Madagascar
rubritarse Pic, 1915; SE Asia
ruficolor Pic, 1915; SE Asia
ruficorne Fairmaire, 1898; Madagascar
rufipenne Pic, 1917; Madagascar
rufipes Pic, 1922; SE Asia
rufitarse Achard, 1920; SE Asia
var. modigliani Pic, 1920
rufofemoratum Pic, 1921; SE Asia
rufonotatum (Pic, 1928)*; SE Asia
rufopygum Lewis, 1893; E Palaearctic
rufum Brancsik, 1893; Madagascar
rugatum Löbl, 1976; New Guinea
salvazai (Pic, 1928)*; SE Asia
sauteri Miwa & Mitono 1943; SE Asia
semiflavum Gestro, 1880; SE Asia
semilimbatum Pic, 1917; SE Asia
septemmaculatum (Pic, 1940)*; SE Asia
septemnotatum Champion, 1927; SE Asia
seriatum Heller, 1917; SE Asia
shelfordi (Achard, 1922)**; SE Asia
shirakii Miwa & Mitono, 1943; SE Asia
shibatai Kimura, 1987; SE Asia
sikorai (Pic, 1915)***; Madagascar
simile Castelnau, 1840; Madagascar
simplicicolle (Pic, 1923)**; SE Asia
sinense Pic, 1954; SE Asia
sinuatum Csiki, 1924; SE Asia
sondaicum Gestro, 1880; SE Asia
var. medionigrum Pic, 1915
var. ruficolor Pic, 1915
striatipenne Gestro, 1880**; SE Asia
striatum Pic, 1920; SE Asia
suarezicum Pic, 1920; Madagascar
subdepressum Pic, 1921++; SE Asia
subelongatum Pic, 1915; SE Asia
subpunctatum (Pic, 1951)*; Madagascar
sulcaticolle (Pic, 1923)**; SE Asia
sulcatum Pic, 1915**; SE Asia
sulcipennes Gestro, 1880**; SE Asia
sylhetense Achard, 1920; SE Asia
takahashii Miwa & Minoto, 1943; SE Asia
takemurai Nakane, 1956; E Palaearctic
testaceum Reitter, 1880; Neotropical
thakali Löbl, 1992; SE Asia
thomasi (Pic, 1926)*; SE Asia
transversale Matthews, 1888; Neotropical
tricolor Achard 1920**; SE Asia
trimaculatum Löbl, 1976; Australia
trinotatum Pic, 1920; SE Asiav
tsushimense Shir=F4zu & Morimoto, 1963; E Palaearctic
tuberculipes (Löbl, 1972)*; SE Asia
undulatum Pic, 1915; Neotropical
unifasciatum Pic, 1916; SE Asia
vagefasciatum Pic, 1920; Madagascar
variabile Matthews, 1888; Neotropical
variegatum Pic, 1915; Neotropical
var. portevini Pic 1920
vernicatum (Pic, 1954); SE Asia
vicinum Pic, 1915; SE Asia
violaceipenne Pic, 1927; SE Asia
viride Löbl, 1978; New Guinea
vitalisi Achard, 1920; SE Asia
vittipenne R. Oberth=FCr, 1884; Neotropical
waterstradti Pic, 1915; SE Asia
yasumatsui Shir=F4zu & Morimoto, 1963; E Palaearctic
yunnanum Fairmaire, 1886; SE Asia
Cerambyciscapha Pic, 1915

dohertyi Pic, 1915; SE Asia
Diatelium Pascoe, 1863

wallacei Pascoe, 1863; SE Asia
spectrum (Vollenhoven, 1865)
var. laterale Achard, 1920

Literature Cited and Annotated Bibliography

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Melsh. and the larva of Scaphium castanipes Kirby with notes on their
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Benick, L. 1952. Pilzk=E4fer und K=E4ferpilze. =D6kologische und statisti= che
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assemblage from Northern India (Coleoptera: Myxomycetes).
Oriental Insects 24:197-218.
Newton A.F., Jr. & M. K. Thayer. 1992. Current classification and
family-group names in Staphyliniformia (Coleoptera). Fieldiana
67:1-92.
Rehfous, M. 1955. Contribution l'=E9tude des insectes des champignons.
Mitt=. Scweiz. Entomol. Ges. 28:1-106.
Scheerpeltz, O. & K. H=F6fler. 1948. K=E4fer und Pilze. Verlag f=FCr
Juge= nd und Volk, Vienna, 351 pp., + 9 pls.

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ACKNOWLEDGMENTS
We thank the Museum d'Histoire naturelle, Geneve for continued support of our project, Rod Hanley for his aid in posting this catalogue, and NSF and Kansas Natural History Museums for support to J.S. Ashe in the development of this Web site.


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This web page was coordinated and by Rodney S. Hanley, University of Kansas Natural History Museum, Division of Entomology (Snow Entomological Research Collection). Last updated: November 1996.

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