Coleoptera
Dytiscidae
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A Classification of the Family Dytiscidae (Coleoptera)
 

Anders N. Nilsson

Department of Biology, BIG, University of Umeâ, S-901 87 Umeâ, Sweden. E-mail: http://www.zo.utexas.edu/faculty/sjasper/beetles/Anders.Nilsson@BIG.UMU.SE

& Robert E. Roughley

Department of Entomology, University of Manitoba, Winnipeg, Manitoba, Canada R3T 2N2.

 

Nilsson et al. (1989) published alpabetical lists of all genus- and family-group names that were known within the family Dytiscidae. As these listings also provided information on the usage of each name in current classification, they also gave this classification, although in a cumbersome way. More recently, Pederzani (1995) rearranged the names into systematic order and published a check list of genus- and family-group taxa of Dytiscidae. This list, however, included several taxonomic judgements not shared by Nilsson et al. (1989). Points of divergence are chiefly due to Pederzani's ambition to conserve well-known names, also when it is contradicted by the International Code of Zoological Nomenclature (ICZN) or results in clearly unnatural (not monophyletic) groupings.

Nilsson & Roughley (1997) have compiled a list of additions and corrections in order to up-date the work of Nilsson et al. (1989). We have also found it useful to produce a systematical list of names that takes advantage of this up-dating and at the same time strictly adheres to the ICZN and excludes taxa that have been shown to be non-monophyletic. Future applications of phylogenetic systematics to more of the dytiscid taxa will no doubt show that a large proportion of the currently recognized taxa are paraphyletic. Until this has been done, we can only view them as arrangements, and continue to use them with this limitation before our eyes.

Only available names are listed. Family group names are given in bold. Names in italics are junior synonyms of the name immediately above them, or in some cases senior synonyms that should or have been suppressed. Subgenera of a particular genus are listed under the appropriate name. Within each category the order of names is alpabetical. An asterisque after a name indicates that it should be conserved.

This list has been published in the newsletter of the Balfour-Browne Club, Latissimus No. 8, 1997. It is here up-dated with the taxa described by Bistrôm et al. (1997) and Spangler (1996). We have now decided to make it available at Sharon Jasperís Water Beetle Page on the Internet. It is our intention to keep it up-dated so that those who want can download the latest version of the list. We also hope that people will mail us their suggestions on additions or corrections so that the list can be improved. In this way it can also serve as the starting-point for a continuous discussion on the systematics of the family Dytiscidae.

 

Literature cited

Bistrôm, O., Nilsson, A.N. & Wewalka, G. 1997. A systematic review of the tribes Hyphydrini Sharp and Pachydrini n. trib. (Coleoptera, Dytiscidae). - Ent. fenn. 8:57-82.

Nilsson, A.N., Roughley, R.E. & Brancucci, M. 1989. A review of the genus- and family-group names of the family Dytiscidae Leach (Coleoptera). - Ent. scand. 20: 287-316.

Nilsson, A.N. & Roughley, R.E. 1997. The genus- and family-group names of the Dytiscidae - additions and corrections (Coleoptera). - Beitr. Ent. 47:359-364.

Pederzani, F. 1995. Keys to the identification of the genera and subgenera of adult Dytiscidae (sensu lato) of the world (Coleoptera Dytiscidae). - Atti Acc. Rov. Agiati, a. 244 (1994), ser. VII, 4(B): 5-83.

Spangler, P.J. 1996. Four new stygobiontic beetles (Coleoptera: Dytiscidae; Noteridae; Elmidae). - Insecta Mundi 10:241-259.

The Classification
Family Dytiscidae Leach, 1815
Subfamily Agabetinae Branden, 1884
Agabetes Crotch, 1873
Subfamily Aubehydrinae Guignot, 1942
Notaticus Zimmermann, 1928
=Aubehydrus Guignot, 1942
Subfamily Colymbetinae Erichson, 1837
Tribe Agabini Thomson, 1867
=Agabinini Crotch, 1873
=Ilybiini Acloque, 1896
Agabinus Crotch, 1873
Agabus Leach, 1817
=Necticus Hope, 1839 (preoccupied)
=Acatodes Thomson, 1859
=Eriglenus Thomson, 1859
=Gaurodytes Thomson, 1859
=Ilybiosoma Crotch, 1873
=Arctodytes Thomson, 1874
=Metronectes Sharp, 1882
=Dichodytes Thomson,1886
=Heteronychus Seidlitz, 1887
=Scytodytes Seidlitz, 1887
=Xanthodytes Seidlitz, 1887
=Apator Semenov, 1899
=Allonychus Zaitzev, 1905
=Asternus Guignot, 1931
=Gabinectes Guignot, 1931
=Agabinectes Guignot, 1932
=Colymbinectes Falkenstrm, 1936
=Nebriogabus Guignot, 1936
=Parasternus Guignot, 1936
=Ranagabus J.Balfour-Browne, 1939
=Dichonectes Guignot, 1945
=Allogabus Guignot, 1951
=Neonecticus Guignot, 1951
=Mesogabus GuÈorguiev, 1969
=Neoplatynectes Vazirani, 1970
Agametrus Sharp, 1882
Andonectes GuÈorguiev, 1971
Carrhydrus Fall, 1922
Hydrotrupes Sharp, 1882
Ilybius Erichson, 1832
=Hyobius Gistel, 1856,
=Ilyobius Gemminger & Harold, 1868
=Idiolybius Gozis, 1886
=Agabidius Seidlitz, 1887
=Ilybidius Guignot, 1948
Leuronectes Sharp, 1882
Platambus Thomson, 1859
Subgenus Agraphis Guignot, 1954
Subgenus Anagabus Jakovlev, 1897
Subgenus Platambus Thomson, 1859
=Stictogabus Guignot, 1948
=Paraplatynectes Vazirani, 1970
Platynectes RÈgimbart, 1879
Subgenus Australonectes GuÈorguiev, 1972
Subgenus Gueorguievtes Vazirani, 1976
=Metaplatynectes GuÈorguiev, 1978
Subgenus Platynectes RÈgimbart, 1878
=Plateocolymbus Gistel, 1857 (suppressed)
=Hypoplatynectes GuÈorguiev, 1972
=Neoplatynectes GuÈorguiev, 1972 (preoccupied)
=Carinonectes Vazirani, 1976
=Notoplatynectes GuÈorguiev, 1978
Tribe Anisomeriini Brinck, 1948
Anisomeria Brinck, 1943
=Anisomera BrullÈ, 1835 (preoccupied)
Senilites Brinck, 1948
Tribe Colymbetini Erichson, 1837
=Cymatopterini Portevin, 1929
=Carabdytini Pederzani, 1995
Bunites Spangler, 1972
Carabdytes Balke, Hendrich & Wewalka, 1992
Colymbetes Clairville, 1806
=Cymatopterus Dejean, 1833
Hoperius Fall, 1927
Meladema Laporte, 1835
=Scutopterus Dejean, 1833 (suppressed)
Melanodytes Seidlitz, 1887
Neoscutopterus J.Balfour-Browne, 1943
=Pseudoscutopterus Hatch, 1953
Rhantus Dejean, 1833
Subgenus Nartus Zaitzev, 1907
=Rantogiton Gozis, 1910
=Ilybiomorphus Porta, 1923
Subgenus Rhantus Dejean, 1833
Tribe Hydronebriini Guignot, 1948
=Hydronebriini Brinck, 1948
Hydronebrius Jakovlev, 1897
Tribe Matini Branden, 1884
Allomatus Mouchamps, 1964
Batrachomatus Clark, 1863
Matus AubÈ, 1836
Subfamily Copelatinae Branden, 1884
=Aglymbinae Branden, 1884
=Lacconectinae Branden, 1884
Agaporomorphus Zimmermann, 1921
Aglymbus Sharp, 1882
Copelatus Erichson, 1832
=Liopterus Dejean, 1833
=Exocelina Broun, 1886
=Pelocatus Zaitzev, 1908
Lacconectus Motschulsky, 1855
=Paralacconectus Vazirani, 1970
Subfamily Coptotominae Branden, 1884
=Coptotominae Brinck, 1948
Coptotomus Say, 1834
Subfamily Dytiscinae Leach, 1815
Tribe Aciliini Thomson, 1867
=Thermonetini Sharp, 1882
=Graphoderini Houlbert, 1934
Acilius Leach, 1817
Subgenus Acilius Leach, 1817
=Heteroelytrus ¡d·m, 1993
Subgenus Homoeolytrus Gobert, 1874
Aethionectes Sharp, 1882
=Afronectes G.M¸ller, 1942
Graphoderus Dejean, 1833
=Graphothorax Motschulsky, 1853
=Prosciastes Gistel, 1856
=Derographus Portevin, 1929
Rhantaticus Sharp, 1882
Sandracottus Sharp, 1882
Thermonectus Dejean, 1833*
Tikoloshanes Omer-Cooper, 1956
Tribe Cybistrini Sharp, 1882
Austrodytes Watts, 1978
Cybister Curtis, 1827
Subgenus Cybister Curtis, 1827
=Trogus Leach, 1817
=Cybisteter Bedel, 1881
=Alocomerus Brinck, 1945
=Megadytoides Brinck, 1945
=Meganectes Brinck, 1945
=Nealocomerus Brinck, 1945
=Gschwendtnerhydrus Brinck, 1945
Subgenus Melanectes Brinck, 1945
Subgenus Scaphinectes ¡d·m, 1993
=Trochalus Dejean, 1833 (preoccupied)
Megadytes Sharp, 1882
Subgenus Bifurcitus Brinck, 1945
Subgenus Megadytes Sharp, 1882
Subgenus Paramegadytes TrÈmouilles & Bachmann, 1980
Subgenus Trifurcitus Brinck, 1945
Onychohydrus Schaum & White, 1847
Subgenus Onychohydrus Schaum & White, 1847
=Homoeodytes RÈgimbart, 1878
Subgenus Sternhydrus Brinck, 1945
Regimbartina Chatanay, 1911
Spencerhydrus Sharp, 1882
Tribe Dytiscini Leach 1815
Dytiscus Linnaeus, 1758
=Leionotus Kirby, 1837
=Macrodytes Thomson, 1859
Hyderodes Hope, 1839
Tribe Eretini Crotch, 1873
Eretes Laporte, 1833
=Eunectes Erichson, 1832 (preoccupied)
=Nogrus Dejean, 1833
Tribe Hydaticini Sharp, 1882
Hydaticus Leach, 1817
Subgenus Guignotites Brinck, 1943
=Isonotus Guignot, 1936 (preoccupied)
Subgenus Hydaticinus Guignot, 1950
Subgenus Hydaticus Leach, 1817
=Icmaleus Gistel, 1856
Subgenus Pleurodytes RÈgimbart, 1899
Prodaticus Sharp, 1882
Subfamily Hydroporinae AubÈ,1836
Tribe Bidessini Sharp, 1882
Africodytes Bistrôm, 1988
Allodessus Guignot, 1953
Anodocheilus Babington, 1841
=Anodontochilus RÈgimbart, 1895
Bidessodes RÈgimbart, 1900
Subgenus Bidessodes RÈgimbart, 1900
Subgenus Hughbosdinius Spangler, 1981
Subgenus Youngulus Spangler, 1981
Bidessonotus RÈgimbart, 1895
Bidessus Sharp, 1882
Boongurrus Larson, 1994
Brachyvatus Zimmermann, 1919
Clypeodytes RÈgimbart, 1894
Subgenus Clypeodytes RÈgimbart, 1894
Subgenus Hypoclypeus Guignot, 1950
=Hypodytes Guignot, 1936 (preoccupied)
Subgenus Paraclypeus Vazirani, 1971
Comaldessus Spangler & Barr, 1995
Geodessus Brancucci, 1979
Gibbidessus Watts, 1978
Hemibidessus Zimmermann, 1921
Huxelhydrus Sharp, 1882
Hydroglyphus Motschulsky, 1853
=Guignotus Houlbert, 1934
Hypodessus Guignot, 1939
=Brachybidessus Gschwendtner, 1954
Leiodytes Guignot, 1936
=Lioclypeus Guignot, 1950
=Liodytes Guignot 1950 (preoccupied)
Limbodessus Guignot, 1939
Liodessus Guignot, 1939
Microdessus Young, 1967
Neobidessus Young, 1967
Neoclypeodytes Young, 1967
Pachynectes RÈgimbart, 1903
Subgenus Pachynectes RÈgimbart, 1903
Subgenus Yoloides Guignot, 1960
Platydytes Bistrôm, 1988
Pseuduvarus Bistrôm, 1988
Sharphydrus Omer-Cooper, 1958
Sinodytes Spangler, 1996
Tepuidessus Spangler, 1981
Trogloguignotus Sanfilippo, 1958
Tyndallhydrus Sharp, 1882
Uvarus Guignot, 1939
Yola Gozis, 1886
=Yolula Guignot, 1950
Yolina Guignot, 1936
Tribe Carabhydrini Watts, 1978
Carabhydrus Watts, 1978
Tribe Hydroporini AubÈ, 1836
=Hydrocoptini Branden, 1884
=Sternopriscini Branden, 1884
=Hygrotini Portevin, 1929
=Siettitiini Smrz, 1982
Antiporus Sharp, 1882
Barretthydrus Lea, 1927
Canthyporus Zimmermann, 1919
Chostonectes Sharp, 1882
Deronectes Sharp, 1882
=Bartheus Houlbert, 1934
Graptodytes Seidlitz, 1887
Haideoporus Young & Longley, 1976
Heroceras Guignot, 1949
Herophydrus Sharp, 1882
Subgenus Dryephorus Guignot, 1949
Subgenus Herophydrus Sharp, 1882
Heterosternuta Strand, 1935
=Heterosternus Zimmermann, 1919 (preoccupied)
=Heterostethus Falkenstrm, 1938 (preoccupied)
Hydroporus Clairville, 1806
=Hydrocoptus Motschulsky, 1853
=Hydatoporus Gistel, 1856
=Sternoporus Falkenstrm, 1930
=Hydroporinus Guignot, 1945
=Hydroporidius Guignot, 1949
Hydrotarsus Falkenstrm, 1938
Hygrotus Stephens, 1828
=Coelambus Thomson, 1860
Hyphoporus Sharp, 1882
Laccornellus Roughley & Wolfe, 1987
Lioporeus Guignot, 1950
=Falloporus Wolfe & Matta, 1981
Megaporus Brinck, 1943
=Macroporus Sharp, 1882 (preoccupied)
Metaporus Guignot, 1945
Nebrioporus RÈgimbart, 1906
Subgenus Nebrioporus RÈgimbart, 1906
=Potamodytes Zimmermann, 1919 (preoccupied)
=Potamonectes Zimmermann, 1921
=Rhabdonectes Houlbert, 1934
Subgenus Zimmermannius Guignot, 1941
=Bistictus Guignot, 1941
Necterosoma MacLeay, 1873
Neoporus Guignot, 1931
=Circinoporus Guignot, 1945
Oreodytes Seidlitz, 1887
=Neonectes J.Balfour-Browne, 1944
=Deuteronectes Guignot, 1945
=Nectoporus Guignot, 1950
Paroster Sharp, 1882
Peschetius Guignot, 1942
Porhydrus Guignot, 1945
Pseudhydrovatus Peschet, 1924
Rhithrodytes Bameul, 1989
Sanfilippodytes Franciscolo, 1979
Scarodytes Gozis, 1914
Siamoporus Spangler, 1996
Siettitia Abeille de Perrin, 1904
Sternopriscus Sharp, 1882
Stictonectes Brinck, 1943
=Stictonotus Zimmermann, 1930 (preoccupied)
Stictotarsus Zimmermann, 1919
=Trichonectes Guignot, 1941
Stygoporus Larson, 1994
Suphrodytes Gozis, 1914
Tiporus Watts, 1985
=Hypodes Watts, 1978 (preoccupied)
Tribe Hydrovatini Sharp, 1882
Hydrovatus Motschulsky, 1853
=Oxynoptilus Schaum, 1868
=Hydatonychus Kolbe, 1883
=Vathydrus Guignot, 1954
Queda Sharp, 1882
Tribe Hyphydrini Sharp, 1882*
=Actobaenini Gistel, 1856
Agnoshydrus Bistrôm, Nilsson & Wewalka, 1997
Allopachria Zimmermann, 1924
=Nipponhydrus Guignot, 1954
Andex Sharp, 1882
Coelhydrus Sharp, 1882
Darwinhydrus Sharp, 1882
Desmopachria Babington, 1841
Subgenus Desmopachria Babington, 1841
Subgenus Hintonella Young, 1981
=Hintonia Young, 1980 (preoccupied)
Subgenus Nectoserrula Guignot, 1949
Subgenus Pachiridis Young, 1980
Subgenus Pachriodesma Guignot, 1949
Subgenus Pachriostrix Guignot, 1950
Subgenus Portmannia Young, 1980
Hovahydrus Bistrôm, 1982
Hydropeplus Sharp, 1882
Hyphovatus Wewalka & Bistrôm, 1994
Hyphydrus Illiger, 1802
=Actobaena Gistel, 1856
=Pachytes Montrouzier, 1860
=Allophydrus Zimmermann, 1930
=Apriophorus Guignot, 1936
=Aulacodytes Guignot, 1936
Microdytes J.Balfour-Browne, 1946
Primospes Sharp, 1882
Tribe Laccornini Wolfe & Roughley, 1990
Laccornis Gozis, 1914
=Agaporus Zimmermann 1919
Tribe Methlini Branden, 1884
=Celinini Branden, 1884
=Celinae Falkenstrm, 1938
Celina AubÈ, 1837
=Hydroporomorpha Babington, 1841.
Methles Sharp, 1882
Tribe Pachydrini Bistrôm, Nilsson & Wewalka, 1997
Heterhydrus Fairmaire, 1869
Pachydrus Sharp, 1882
Tribe Vatellini Sharp, 1882
Derovatellus Sharp, 1882
Subgenus Derovatellus Sharp, 1882
Subgenus Varodetellus Bistrôm, 1979
Macrovatellus Sharp, 1882
=Platydessus Guignot, 1955
Mesovatellus TrÈmouilles, 1995
Vatellus AubÈ, 1837
=Leucorea Laporte, 1835 (suppressed).
Subfamily Hydroporinae Incertae sedis
Amarodytes RÈgimbart, 1900
Hydrodessus J.Balfour-Browne, 1953
=Brinckius Guignot, 1957
Kuschelydrus Ordish, 1976
Morimotoa UÈno, 1957
Phreatodessus Ordish, 1976
Terradessus Watts, 1982
Typhlodessus Brancucci, 1985
Subfamily Laccophilinae Gistel, 1856
Africophilus Guignot, 1947
Australphilus Watts, 1978
Japanolaccophilus SatÙ, 1972
Laccodytes RÈgimbart, 1895
Laccophilus Leach, 1817
Laccoporus J.Balfour-Browne, 1939
Laccosternus Brancucci, 1983
Napodytes Steiner, 1981
Neptosternus Sharp, 1882
Philacus Guignot, 1937
Subgenus Philacilus Guignot, 1937
Subgenus Philacus Guignot, 1937
Philodytes J.Balfour-Browne, 1939
Subfamily Lancetinae Branden, 1884
Lancetes Sharp, 1882

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